Martin Spies


By Dipl.-Biol. Martin Spies

A) In printed media

1. Spies, M., E. J. Fittkau & F. Reiss 1994. The adult males of Parachironomus Lenz, 1921, from the Neotropical faunal region (Insecta, Diptera, Chironomidae). – Spixiana Suppl. 20: 61-98.

2. Spies, M. 1996. Neotropical bibliography and catalog: announcement of work in progress and call for contributions. – Chironomus 9: 13-20.

3. Spies, M. & F. Reiss 1996. Catalog and bibliography of Neotropical and Mexican Chironomidae (Insecta, Diptera). – Spixiana Suppl. 22: 61-119.

4. Gonser, T. & M. Spies 1997. Southern hemisphere Symbiocladius (Diptera, Chironomidae) and their mayfly hosts (Ephemeroptera, Leptophlebiidae). Pp. 455-466 in: Landolt, P. & M. Sartori (eds), Ephemeroptera & Plecoptera. Biology – ecology – systematics; MTL, Fribourg (Switzerland), XI+569 pp.

5. Spies, M. 1998a. Three species of Tanytarsus involved in California midge nuisance problems: descriptions, ecology, and faunal relations (Insecta, Diptera, Chironomidae). – Spixiana 21: 253-270.

6. Spies, M. 1998b. Resolution of Tanytarsus mendax Kieffer, 1925, T. dibranchius Kieffer in Zavrel, 1926, and associated names (Diptera: Chironomidae). – Studia dipterol. 5: 195-210.

7. Spies, M. (1998) 1999a. Cricotopus (Isocladius) subletteorum, a new species of Chironomidae (Diptera) from the southwestern United States. – J. Kansas Ent. Soc. 71(3): 199-207.

8. Fittkau, E. J. & M. Spies 1999. Obituary notice. In memory of Friedrich Reiss (24 December 1937 – 17 August 1999). – Chironomus 12: 17-23.

9. Spies, M. 1999b. Last minute news. Overlooked historic descriptions. – Chironomus 12: 24. (siehe Spies1999.pdf)

10. Cranston, P. S. & M. Spies (1999) 2000. The immature stages and phylogenetic position of Nilodosis Kieffer, an Afrotropical genus of Chironomini (Diptera: Chironomidae). – African Ent. 7: 249-259. (siehe nilodosis.pdf)

11. Andersen, T., A. Contreras-Ramos & M. Spies 2000. Chironomidae (Diptera). Pp. 581-591 in: Llorente B., J., E. González S. & N. Papavero (eds), Biodiversidad, taxonomía y biogeografía de artrópodos de México: hacia una síntesis de su conocimiento. Vol. II; Universidad Nacional Autónoma de México, Mexico City.

12. Spies, M. 2000a. Chironomidae. In: Ziegler, J. & F. Menzel (eds), Die historische Dipteren-Sammlung Carl Friedrich Ketel. Revision einer zwischen 1884 und 1903 angelegten Sammlung von Zweiflüglern (Diptera) aus Mecklenburg-Vorpommern. – Nova Suppl. Entomol. 14: 44-45.

13. Baehr, M. & M. Spies (eds) 2000. Contributions to chironomid research in memory of Dr. Friedrich Reiss. – Spixiana 23(2): 97-224 and 23(3): 225-274.

14. Spies, M. 2000b. A contribution to the knowledge of Holarctic Parachironomus Lenz (Diptera: Chironomidae), with two new species and a provisional key to Nearctic adult males. – Tijdschr. Ent. 143: 125-143.

15. Spies, M. 2000c. Non-biting ‘nuisance‘ midges (Diptera: Chironomidae) in urban southern California, with notes on taxonomy, ecology and zoogeography. Pp. 621-628 in: Hoffrichter, O. (ed.), Late 20th century research on Chironomidae: an anthology from the 13th International Symposium on Chironomidae; Shaker, Aachen.

16. Spies, M. 2001. When is a nomen really dubium? Toward real stability in chironomid taxonomy through better symbiosis with the classic collections. – Chironomus 14: 7-10. Link zum PDF

17. Spies, M., J. E. Sublette, M. F. Sublette, W. F. Wülker, J. Martin, A. Hille, M. A. Miller & K. Witt 2002. Pan-American Chironomus calligraphus Goeldi, 1905 (Diptera, Chironomidae): species or complex? Evidence from morphology, karyology and DNA sequencing. – Aquatic Insects 24(2): 91-113.

18. Spies, M. 2002. Professor Ernst Josef Fittkau – 75 years, 50 years for chironomid research. – Chironomus 15: 2-13. Link zum PDF

19. Michiels, S. & M. Spies 2002. Description of Conchapelopia hittmairorum, spec. nov., and redefinition of similar western Palaearctic species (Insecta, Diptera, Chironomidae, Tanypodinae). – Spixiana 25(3): 251-272.

20. Foley, I. A., M. A. Ivie & M. Spies (2003) 2004. Pachystigmus Hellén, 1927, a substitute name for Noserus Foerster, 1863 (Hymenoptera: Braconidae), not Noserus LeConte, 1862 (Coleoptera: Zopheridae). – Insecta Mundi 17(1-2): 103-106.

21. Spies, M. & O. A. Sæther 2004. Notes and recommendations on taxonomy and nomenclature of Chironomidae (Diptera). – Zootaxa 752: 1-90. Link zum PDF

22.  Spies, M. 2005. On selected family-group names in Chironomidae (Insecta, Diptera), and related nomenclature. – Zootaxa 894: 1-12. Link zum PDF

23. Spies, M. 2005. An index to selected scientific names in Edwards’ (1929) monograph on British Chironomidae. – Chironomus 18: 11-24. Link zum PDF

24. Spies, M. 2006. Case 3344. Pseudorthocladius Goetghebuer, 1943 and Mesosmittia Brundin, 1956 (Insecta, Diptera, CHIRONOMIDAE): proposed conservation of the generic names. – Bull. Zool. Nomenclature 63(2): 122-126. Link zum PDF

25. Spies, M. 2007. Case 3355. ORTHOCLADIINAE Kieffer, 1911 and Orthocladius van der Wulp, 1874 (Insecta, Diptera, CHIRONOMIDAE): proposed conservation of subfamilial name and fixation of type species. – Bull. Zool. Nomenclature 64(1): 45-53. Link zum PDF(PDF is unofficial, corrected version)

26. Spies, M. & N. L. Evenhuis 2007. [Comment] (1). Pp. 124-125 in: Comments on the proposed conservation of the subfamilial name ORTHOCLADIINAE Kieffer, 1911 and on the proposed type-species fixation for Orthocladius van der Wulp, 1874 (Diptera, CHIRONOMIDAE). (Case 3355; see BZN 64: 45-53). – Bull. Zool. Nomenclature 64(2): 124-126. Link zum PDF

27. Spies, M. 2008. Parachironomus valdiviensis, spec. nov., and other changes to nomenclature of Neotropical Chironomidae (Insecta, Diptera). – Spixiana 31: 173-175.


28. Vinogradova, E.M., Riss, H.W. & M. Spies 2009. New species of Tanytarsus van der Wulp, 1874 (Diptera: Chironomidae) from Central America. – Aquatic Insects 31(1): 11-17.

29. Spies, M., Andersen, T., Epler, J.H., Watson, C.N., Jr. 2009. Chironomidae (non-biting midges). Pp. 437-480 in: Brown, B.V. et al. (eds.) Manual of Central American Diptera, vol. 1. – NRC Research Press, Ottawa.

30. Orendt, C. & M. Spies 2010. Bestimmungsschlüssel Chironomini: Illustrierter Bestimmungsschlüssel für die mitteleuropäischen Larven der Zuckmücken-Unterfamilien (Diptera: Chironomidae) sowie der Unterfamilie Chironominae, Tribus Chironomini. Unter Verwendung vorwiegend makroskopischer Merkmale. – Leipzig, 59 pp.

31. Stur, E. & M. Spies 2011. Description of Chaetocladius longivirgatus sp. n., with a review of C. suecicus (Kieffer) (Diptera: Chironomidae). – Zootaxa 2762: 37-48. PDF…

32. Ashe, P., Spies, M. 2011. A chronological bibliography, with dates of publication, of J. J. Kieffer’s works on Chironomidae (Diptera). Pp. 3-14 in: Wang, X., Liu, W. (eds.) Contemporary chironomid studies. Proceedings of the 17th International Symposium on Chironomidae. – Nankai University Press, Tianjin, China.

33. Spies, M. 2011. Typical types – a swan song? Observations on chironomids in the Linnean collections, and corresponding general considerations. – Chironomus 24: 30-35. [s.]

34. Kotrba, M. & Spies, M. 2011. Obituary for Wolfgang Schacht (1939-2011). – Spixiana 34: 146.

35. Orendt, C. & M. Spies 2012. Chironomini (Diptera: Chironomidae). Keys to Central European larvae using mainly macroscopic characters. 2nd, revised edition. – Leipzig, 64 p., 450 illustrations. ISBN 978-3-00-038842-2.

36. Orendt, C. & M. Spies 2012. Chironomus (Meigen) (Diptera: Chironomidae). Key to the larvae of importance to biological water analysis in Germany and adjacent areas. Bilingual edition (German/English). – Leipzig, 24 p., 47 illustrations. ISBN 978-3-00-038789-0.

37. Gilka, W. & M. Spies 2012. Neotype designation for Cladotanytarsus (Lenziella) bicornutus (Kieffer), and first description of the distinctive female (Diptera: Chironomidae: Tanytarsini). – Zootaxa 3545: 83-88.

38. Hausmann, A., Spies, M. & J. Diller 2012. In memoriam Prof. Dr. Ernst Josef Fittkau (22.07.1927 – 12.05.2012). – Spixiana 35(2): 161-176. l to pdf

39. Spies, M. & S. J. Bolton 2013. On the first record from Britain of Parachironomus elodeae (Townes) Diptera, Chironomidae). – Dipterists Digest 20(1): 79-85.

40. Jäger-Zürn, I., Spies, M., Philbrick, C. T., Bove, C. P. & A. Mora-Olivo 2013. Plant galls (cecidia) in the neotropical water plant family Podostemaceae induced by larvae of Chironomidae (Diptera). – Spixiana 36(1): 97-112.

41. Spies, M. 2013a. A simple method for slide-mounting chironomid hypopygia in lateral view (and for similar preparations). – Chironomus 26: 47-48.

42. Spies, M. 2013b. A survey of publications with chironomid content represented in bibliographic databases on the web. – Chironomus 26: 56-57.

43. Bello González, O. C., Spies, M., Téllez Martínez, B. 2013. Estado del conocimiento de la familia Chironomidae (Insecta: Diptera) en Cuba. – Dugesiana 20(2): 233-242.

44. Haszprunar, G., Spies, M. 2014. An integrative approach to the taxonomy of the crown-of-thorns starfish species group (Asteroidea: Acanthaster): A review of names and comparison to recent molecular data. – Zootaxa 3841(2): 271-284.

45. Orendt, C., Spies, M. 2014. Chironomus (Meigen) (Diptera: Chironomidae). Bestimmungsschlüssel zu den für die biologische Gewässeranalyse bedeutenden Larven in Deutschland und angrenzenden Gebieten. Nachdruck mit kleineren Korrekturen. / Chironomus (Meigen) (Diptera: Chironomidae). Key to the larvae of importance to biological water analysis in Germany and adjacent areas. Reprint with minor corrections. – Published by the authors; Leipzig. ISBN 978-3-00-038789-0.

46. Orendt, C., Dettinger-Klemm, A., Spies, M. 2015. Bestimmungsschlüssel für die Larven der Chironomidae (Diptera) der Brackgewässer Deutschlands und angrenzender Gebiete. In: Schilling, P. (ed.): Bund/Länder-Messprogramm für die Meeresumwelt von Nord- und Ostsee (BLMP) – Ber. Qual.sich.stelle 2013/1. Deutschland/Umweltbundesamt, Dessau-Roßlau, etc.; 242 pp. [“2013”]

47. Spies, M., Dettinger-Klemm, A. 2015. Diagnoses for Nubensia, n. gen. (Diptera, Chironomidae, Chironomini), with the first full descriptions of the adult female and larva of N. nubens (Edwards, 1929). – Zootaxa 3994(1): 109-121.

48. Cranston, P. S., Martin, J., Spies, M. 2016. Cryptic species in the nuisance midge Polypedilum nubifer (Skuse) (Diptera: Chironomidae) and the status of Tripedilum Kieffer. – Zootaxa 4079(4): 429-447.

49. Cranston, P. S., Martin, J., Mulder, M., Spies, M. 2016. Clarification of Einfeldia Kieffer, 1922 (Diptera: Chironomidae) with E. australiensis (Freeman, 1961), comb. n. based on immature stages. – Zootaxa 4158(4): 491-506.

B) In electronic media

1. Spies, M. (authorship explicitly stated in the product and related documentation) 1997. Corrections and additions to data on the Chironomidae – In: Poole, R. W. & P. Gentili: Nomina Insecta Nearctica. A database application for all the orders; CD-ROM; Entomological Information Services, Rockville, Maryland.

2. Spies, M. 1999. A listing of Chironomidae reported from California. Internet page

3. Spies, M. 1999. A listing of Ceratopogonidae reported from California. Internet page

4. Spies, M. 1999-2003. Friedrich Reiss (24.12.1937 – 17.8.1999). Internet page; no longer available.

5. Spies, M. 1999. List of papers on Chironomidae published in SPIXIANA. Internet page

6. Sæther, O. A. & M. Spies 2004-2011. Fauna Europaea: Chironomidae in: de Jong, H. (ed.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 1.2. Internet database, outdated version.

7. Brabec, K., Janeček, B. F. U., Rossaro, B., Spies, M., Bitušík, P., Syrovátka, V. & A. Schmidt-Kloiber 2007. Chironomidae Indicator Database in: Euro-limpacs project, Workpackage 7 —Indicators of ecosystem health, Task 4, version 5.0. Internet database

8. Sæther, O. A. & M. Spies 2011-2012. Fauna Europaea: Chironomidae in: Beuk, P. & T. Pape (eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.4. Internet database, outdated version.

9. Aagaard, K., Hoffrichter, O., Spies, M., Daverdin, M. & T. Ekrem 2012-present. An online bibliography of the Chironomidae. NTNU Museum of Natural History and Archaeology, Trondheim, Norway. Internet database

10. Sæther, O. A. & M. Spies 2012-2013. Fauna Europaea: Chironomidae in: Beuk, P. & T. Pape (eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.5. Internet database, outdated version.

11. Sæther, O. A. & M. Spies 2013. Fauna Europaea: Chironomidae in: Beuk, P. & T. Pape (eds.) Fauna Europaea: Diptera Nematocera. Fauna Europaea version 2.6. Internet database, online since April 2013.